Contributions in Phycology
This is the cover of the book which Bruce and I published to raise funds for the Bold Award

        On August 29, 1967, Dr. Harold C. Bold and his wife, Mary,
attended a banquet held in his honor at Caldwell near Texas
A & M University, College Station, Texas. Approximately 25 of
his former graduate students accompanied him, some having
traveled several thousand miles to share that memorable evening
with their esteemed major professor.
        Words cannot convey the personal experiences recalled by
each of us that evening which had led to our very close friend-
ship with Dr. Bold. Some of us were first drawn to him by the
magnetism of his lectures on plants, wherein organisms and their
parts seemed to come alive from a combination of chalk-on-black-
board finesse and humorous stories and anecdotes spanning the
many years of Dr. Bold's professional botanical experience.
        Others of us may have recalled the deep interest he had in each
of our lives, an interest which continued beyond the degree. Dr.
Bold instilled self-confidence in some of us and provided oppor-
tunities for us to pursue more rapidly the paths to our post-
doctoral goals.
        Most of us will recall with amusement the numerous times he
approached a colleague or one of us asking apologetically and
humbly for a lucid explanation of some scientific concept or
observation, only to discover that he could explain it more
thoroughly than the person he was asking. Again, we may smile
when during our own lectures, we attach one of his favorite quo-
tations or paraphrases of them, to an appropriate plant life-cycle,
such as:

"Many are called, but one is chosen"1


"Of those that run in the race, only one (usually) receiveth the prize." 2

        And similarly, when confronted with a taxonomic dilemma, we
may repeat an expression which goes back to Dr. Bold's major
professor, the late Dr. Robert A. Harper and perhaps even to
Harper's major professor, the renowned Dr. Eduard Strasburger:
"Nature mocks at human categories."
1 In reference to the number of juvenile sporophytes which develop from
more than one zygote on a single gametophyte.
2 In reference to polyembryony and seed development in conifers.

        Before the banquet at Caldwell ended, Dr. Bold rose to
recount humorous events in the predoctoral lives of each of his
former students in attendance. It was then and subsequently
that the idea for this small book congealed--a book to contain
contributions of a select number of Dr. Bold's former students
thereby reflecting a portion of his impact, as a great teacher,
upon the botanical world. This book, however humble, is sym-
bolic of the inspiration given by him to us and to many others.

Resume of Harold C. Bold

HAROLD C. BOLD was born in New York City, June 16, 1909.
He received his B.A. degree from Columbia University in 1929,
his M.S. from the University of Vermont in 1931, and his Ph.D.
from Columbia in 1933. Positions held by Dr. Bold include:

Instructor of Botany, University of Vermont (1929-31).

Instructor of Biology, Vanderbilt University (1931-9).

Visiting Lecturer (1939-40) and Assistant Professor of
Botany, Barnard College, Columbia University (1940-2);
U.S. Naval Reserve, active duty (1942-5);

Associate Professor (1946-9) and Professor of Biology,
Vanderbilt University (1949-57).

Professor of Botany, University of Texas (1957-);

Chairman, Dept, of Botany, University of Texas (1962-7);

Chairman, Division of Biological Science, University of
Texas (1967-).

Dr. Bold also has held a number of summer appointments at
such institutions as the Chesapeake Biological Laboratory
(1934-7) and the Marine Biological Laboratory, Woods Hole,
Massachusetts (1954-8). He has served as Secretary (1955-8),
Vice-President (1965), and President (1968) of the Botanical
Society of America, Editor-in-Chief of the American Journal of
Botany (1958-65), and President of the Phycological Society of
America (1955-6).



        This book has at least one unique mission: It is destined to serve as a means
by which we (the students of Harold Bold) hope to establish one or more annual
awards to encourage outstanding work in the field of phycology, especially the
aspect of teaching. In view of this mission and the book's dedication to Harold
Bold by his former students, this volume contains exclusively original contribu-
tions by authors who were exposed, at some time in the past, to Harold Bold's
teaching and/or research.
        The editors and authors of this small book would be pretentious even to suggest
that the contents herein represents a cross section of the broad area encompassed
by Bold's 60 or more Ph.D. students. Indeed, a number of these students are not
phycologists, but specialize in other areas of botanical or microbiological study.
Neither does this book cover any single area of phycology intensively. Rather,
the reader will find here a variety of topics, including reports of original research
and a few essays, with generally similar formats but written in the individual
author's styles, and for the purpose of informing the student, teacher, and re-
searcher of algae of current trends in the modern field of phycology.
        The struggle to publish this book has not been without trauma. The book
went to press in August 1970, and passed through the hands of two printers
before the editors were satisfied with the quality of reproduction of photo- and
electron-micrographs. As a result of these delays, we apologize humbly for a few
reference citations, addresses and statements in the bibliographic section, and
occasional statements which may deserve some up-dating.
        All profits from the sale of this book will go toward establishing the one or
more awards referred to above. In achieving this goal, we shall remain perpet-
ually grateful to a number of Harold Bold's former students for their assistance
in financing the publication of this book. They are: Gina Arce, Pat Archibald,
Emily Bell, Harry Bischoff, Malcolm Brown, Neal Buffaloe, Srisumon Chantana-
chat, Elenor Cox, Temd Deason, Richard Dykstra, Peter Edwards, Robert Groover,
Waiter Herndon, Larry Hoffman, Don Kapraun, Norma Lang, Bob McLean, Karl
Matter, Jerome Metzner, Rev. John Mullahy, Bruce Parker, George Pessoney,
Takashi Sawa, Eugene Shen, Richard Smith, Richard Starr, Dempsey Thomas,
Frank Trainer, Pat Walne.

Table of Contents
Contributions in Phycology
Part I. Systematics and Phylogeny

Borodinellopsis texensis: Gen. et sp. nov. A new alga from the

A circumscription of the genus Protosiphon, BY DEMPSEY L. THOMAS

Stichococcus seguoieti sp. nov., BY GINA ARCE

Nutritional attributes as taxonomic tools in phycology, BY RICHARD L. SMITH

On the evolution of isogamy to oogamy, BY BRUCE C. PARKER

Part II. Morphology and Cytology

The effect of aging on thylakoid configuration and granular inclusions in

Sexual reproduction in Volvox africanus, BY RICHARD C. STARR

The male initial and mitosis in Volvox, BY TEMD R. DEASON AND

Development of, form in Scenedesmus, BY FRANCIS R. TRAINOR

Observations on the fine structure of Oedogonium. VII. The oogonium
prior to fertilization, BY LARRY R. HOFFMAN

Comparative ultrastructure of eyespots in selected euglenoid
flagellates, BY PATRICIA L. WALNE

The ultrastructure of the theca of the marine dinoflagellate, Ensiculifera
loeblichii sp. nov, BY ELENOR R. COX and HOWARD J. ARNOTT

Part III. Ecology and Physiology

Zoosporogenesis and resistant-cell formation in Hormidium flaccidum,

Formation and resistance of akinetes of Zygnema, BY ROBERT J. MCLEAN

Cultivation of Chara in defined medium, BY EUGENE Y. F. SHEN

Effects of light intensity, daylength, and temperature on growth and
reproduction of Calithamnion byssoides, BY PETER EDWARDS

Studies of Hawaiian fresh-water and soil algae. I. The atmospheric
dispersal of algae and fern spores across the island of Oahu,

O.i.n.a. = other information not available.
(needs lots of careful editing as the scan was not good on this section! please forgive!)
ARCE, GINA--b. Cuba, 1929; Dept. of Botany, Fresno State College, Fresno, Calif
Education-B.A.., George Peabody College, 1949; M.A., 1950; Ph.D., Vanderbilt Univer-
sity, 1956.
Academic Experience- Prof.. of Botany, Fresno State College.
Dissertation-"Some Chlorophyceae from Cuban Soils."
Interests-Teaching, general botany and plant morphology; research, algae of the Moiave

ARCHIBALD, PATRICIA-b.. Olney, Ill., 1934; Dept. of Biology, Slippery Rock College, Slippery
Rock, Pa.
Education--B.S., M.A., Ball State University, 1961; Ph.D., University of Texas, Austin,
Academic Experience-Instructor, Elkhart' Senior High School, Elkhart, Ind. 1959-6~;
instructor, Palm Beach Jr. College, Lake Worth, Florida, 19~i4~6; instructor, Kuyahoga Com-
munity College, Cleveland, Ohio, 196~-67; Assistant Prof., Slippery Pock College, Slippery
Rock, Pa., 19~9-
Dissertdiorz-"A Study of Bog Algal Flora with Special Reference to the Genus Chloro-
Interests-Teaching phycoaogy, morphology; research, algae, particularly of bogs, algae of
acid mine polluted streams, Latin America Chlolrococcales.

BAKER, AILSIE FAYE-b. Memphis, Term., 1944; Dept. of Biological Sciences, Chicol State Col-
lege, Chico, Calif.
EdUCd~07t--B.A., Univ. of Tenn., 196~; Ph.D., Univ. of Texas, Austin, 1969.
Academic Expe7·ielz~:e-·Assistal~:t Prof., Dept. of Biological Sciences, Chico State College,
Chico, Calif.
Dissertation-"Tax~nor~lic Studies in the Oscillatoriaceae."

BELL, EMLLY J.-b. Chattan~K>ga, Tenn., June 20, 19·27; Dept. of Biological Sciences, Univ. of
Cincinnati, Cincinnati, Ohio.
Education--B.A., Wesleyan College, 1948; M.A., Vanderbilt Univ., 19~9; Ph.D., Vanderbnt
Univ., 1953.
Academic Experi~nce-Instructor in Biology, Vanderbilt Univ., 1951-52; Instructor in Baci
teriology, Univ. of Keautucky, 1952c53; Instructor in Biocher~)listry, Vanderbilt Univ. School of
Medicine, 19~F~i355; Assistant Prof. of B~ochre~kistry, Vanderbilt Univ. School of Medicine, 1.955-
57; Assistant Prof. of Bacteriology, Univ. of Cincinnati, 1957~0; Associate Prd. of Bacteriolo:gy,
Univ. of Cinc3hnati, 1961-67; Prof. of Bacteriology, Univ. of Cinciz~ati, 19~1-
Honors and Azliards-Post-doctoIral, Biology Division, Oak Ridge National Laboratory,
195;P55~; Interamerican Fellow for training in Tropical Medicine, N.I.H., July 19~1; Inter-
american Research Fellow, Caracas, VenezueZa, 1963; Interamerican Research Fellow, Caribbean
area, summer 19~7; Invited participant, First International Congress of Parasitology, Rome,
Italy, 1964; Diplomate in Public Health and Medical Laboratozy Bacteriology from the American
Academy of Microbiology.
Dissertationc"A Physiological Study of Chlamydomofir~s ch2rrmydogarmr Bold" (also with
Dr. Ilda McVeigh).
Interests-Teaching, bacteriology, ~immunology; research, micro;bial physiology with em-
phasis on the metabolic regulation of intermediary metabolism.

BISCHOFF, HARRY W.-b. Evansville, Ind., May 15, 19~22; Dept. of Biology; Texas Lutheran
College, Seguirq Texas.
Educatio~-B.A., Evansville Univ., 19·Q~; M.S., Vanderbilt Univ., 1952; Ph.D., Univ. of
Texas, Austin; 1963.
Parker and Brown
Academic Experienc~LPro·f. of Biology and Chairman, Dept. of Biology, Texas Lutheran
College, 1950-
Honors and Award~--NSF Faculty Fellowship, Austin, Texas, 1959; NSF Academic Year
Extension Grant, 1983, 19~4.
Dissertatio~c"I. The Soil Algal Flora of En~hanted Rock, II. Thorea rieket sp. nov. and
Related Species."
Interests--Teaching, biology; research, fresh-water, soil, and marine algae, aquatic liver-
worts and phytoplankton.

BROWN, JR., RICHARD MACOLM-b. Pampa, ?'exas, Jan. 2, 1939; Dept. of Botany, Univ. of
North Carolina, Chael Hill, N. C.
Educatiot~--B.A., Univ. of Texas, Austin, 19~1; Ph.D., 1964.
Academic Experience-Assistant Prd. of Botany, Univ. of Texas, Austin, 1965-8&; Associate
Prof. of Botany, Univ. of N.C., 196·8-
Honors and Awards--NSF Undergraduate Research Fellow, summers, 1960161; Post-
doctoral Research, Univ. of Hawaii, Honolulu, summers, 1964-65, Univ. of Texas, 1984-65;
NSF Post-doctoral Fellow, Botanisches Institut der Universitat, Freiburg, West Germany, 1968-
Dissertation-"Comparative Studies of the Algal Genera Tetmcystis and Clzlorocorcum."
It~erests--Teaching, fresh-water phycology, gen~eo~al biology, plant cytology, plant ultra-
structure; research, algal viruses, air-borne algae, Golgi apparatus and cell wall formation,
lichen ultrastructure, pyrenoid structure and function, systematics, morphology, and ulh-a-
structure of chrysophycean algae, biochemical systematics of cklorophycean algae, scientific
research and teaching films of algae.

BUFFALOE, NEAL DOLLISON-b. Leachville, Ark., Nov. 15, 1924; Dept. of Biological Science,
State College of Ark., Conway, Ark.
Educc~ion--B.S., David Lipscomb College, 1949; M.S., Vanderbilt Univ., 1952; Ph.D., 1957.
Academic Erperienc~--Prof. of Biology and Chairman, State College of Ark., 1957-
HonM·s and Awards--President, Ark. Academy of Science, 1959-60.
Dissertation--"A Cytological Study of Four Species of Chlamydomonas."
Interests--Teaching, microbiology, genetics, history and philosophy of biological science;
research, cytology of algae, writing of textbooks in the biological sciences.

CAIN, BROTHER JOSEPH R.-b. Indianapolis, Ind., April 13, 1933; I)ept, of Biology, St. Edwards
Univ., Austin, Texas.
Education--B.S., St. Edwards Univ., 1955; M.S., Univ. of Notre Dame, 1961; Ph.D., Univ.
of Texas, Austin, 1963.
Academic Experience--Prof, of Biology, St. Edwards Univ., 1963-
Disse7tatiotc"The Morphology, Taxonomy, and Physiology of Thirty-eight Chlamydorn-
onas-like Algae."
Interests--Teaching, cryptogamic botany; research, biology of Chlamydomonas and other
fresh-water chlorophycean algae, m~rphology of various cryptogar~s from central Texas.

CHANTANACHAT, MISS SRISUMON-b. Bangkok, Thailand, March 23, 1933; Dept. of Botany,
Faculty of Sciences, Chulalongkorn Univ., Bangkok, Thailand.
Educaticm--B.S., Chulalongkorn Univ., 1956; M.A., Univ. of Texas, Austin, 1962.
Academic Experience--Senior Lecturer, Dept. of Botany, Faculty of Sciences, Chulalong-
korn Univ., Bangkok, Thailand, 1~2·
Thesis--"An Investigation of the Algae in Desert Soils."
mterests--TTeaching, phycology.

COX, ELENOR RAY-b. Georgetown, Texas, Nov. 4, 1930; Dept. of Biology, Texas A&M Univ.,
College Station, Texas.
Edzrcation--B.A., Rice Univ., 1952; M.A., Univ. of Texas, Austin, 1961; Ph.D., 1968.
Academic Experi~~ce-Assistant Prof., Dept. of Biology, Texas A&M Univ., 1966-
ContribuZions in Phycology
Dissertdion-"Taxonomic, Morphological and Physiological Studies of the Algal Genus
InterestJLTaaching, fresh-water and marine phyoo;logy; research, fresh-water and marine
dinoflagellates, epiphytic algae, algal ultrastructure, systematics and morphology.

DEASON, TEMD ROBERT-b. York, Ale., Oct. 13, 1931; Dept. of Biological Science; Univ. of
Ala., University, Ala.
Educdiott--TE.S., Univ. of Ale., 1454; M.S., 1958; Ph.D., Univ. of Texas, Austin, 1960.
Academic E~perience-Assistant Prof. of Biological Science, Univ. of Delaware, 1960-6~1;
Assistant Prof. of Biology, Univ. of Ala., 1961-63; Associate Prof. of Biology, Univ. of Ala.,
Honors and Azoar~--Secretary, Ala. Chapter of the Sigma Xi, 196&
Dissertation-''Exploratory Studies of Texas Soil Algae."
interests-Teaching and research, morphology and taxonomy of fresh-water algae, algal

DYKSTRA, RICHARD F.-b. HU11, Iowa, April 17, 1940; Uni~. Texas, Austin.
Edz~cation--B.S., Iowa State Univ., 1963; M.A., Univ. of Texas, Austin, 1966.
Honors crmd Awards-Society of Sigma Xi.
Dissertation--"An Investigation of Some Algae of the Texas Gulf Coast."
Interests--Teaching, general botany, general biology, phycology; research, morphology and
physiology of algae.

EDWARDS, PETER-b. Chorley, Lancashire, England, July 4, 1943; Dept. of Botany, Univ. of
Durham, England.
Education--B.S. (biology), Univ. of Liverpool, England, 1965; B.S. (botany), Univ. of
Liverpool, Englanld, 19~6~; Ph.D., Unjv. Texas, Austin, 1969.
Academic Experience-Post-doctoral Fellowship, Dept. of Bot., Univ. of Durham, England,
Dissertation--"Field and Cultural Studies of the Seasonal Periodicity of Growth and
Reproduction of Selected Texas Marine Algae."
Interests-Experimental marine, algal ecology, bryology.

GROOVER, ROBERT DON-b. Harts~lle, Ala., April 29, 1938; Dept. of Biology, Tulanle Univ.,
New Orleans, La.
Education--B.S., Univ. of Ale., 1960; M.S., 19~6~; Ph.D., Univ. of Texas, Austin, 1908.
Academic Experience-Instructor of Biology, Ala. College, 1963-86; Assistant Prof. of
Biology, Tulane Univ., 1968-
Dissertation-'%xperimental Studies on Chlorosphaeracean and other Edaphic AIgae."
Interests--Teaching, phycology, plant morphology, general biology; research, morphology
and taxonomy of algae.

HARRIS, W. BRYCE-McCauley School, Chattanooga, Tenn. o.i.n.a.

HERNDON, WALTER R.-Dept. of Botany, Univ. of Tenn., Knoxville, Tenn. o.i.n.a.

HOFFMAN, LARRY RONALD-b. Sigourney, Iowa, May 12, 1936; Dept. of Botany, Univ. of Ill.,
Urbana, Ill.
Education--B.S., Iowa State Univ., 1958; Ph.D., Univ. of Texas, Austin, 1961.
Academic Experience-Assistant Plof. of Botany, Univ. of Ill., 1982-68; Associate Prof.
of Botany, Univ. of Ill., 196&  ; sabbatic leave, Univ. of Leeds, England, 1969-70.
Honors and Atoards--NSF Post-doctoral Fellow, Univ. of Leeds, England, 1961-62.
Dissertatioll~-"Studies on the Morphology, Cytology and Reproduction of Oedogazium and
Interests--Teaching, general botany, plant morphology, phycology; research, algal. fine
structure, with special emphasis on basal bodies, flagella, and flagellar appendages, sexuality
in algae, morphology, cytology and reproduction in the Oedogoniales.
192                         Parker and Brown


JOHNSTON, LAVERNE ALBERT-b. Hallettsville, Texas, March 7, 1930; Dept. of Botany, Univ.
of Texas, Austin, Texas.
Education--B.A., Baylor Univ., 1951; M.R.E. (religious education), Southwestern Baptist
Theological Seminary, 1954; M.A., Baylor Univ., 1957; Predoctoral studies, Univ. of Texas,
Austin, 19~j0-62 (with H. C. Bold).
Academic Experience-Assistant Prof. of Biology, Baylor Univ., 1957-60.
Honors and Azoards--NSF Teacher-Research Fellowship, 1960.
Dissertation--( Incomplete)  "A Systematic Study of Algae Isolated from Central Texas
Interests--(with Marshall C. Johnston) monograph of the Rhamnaceae of the world.

KANTZ, JR., PAUL THOMAS-b. Jacksonville, Texas, Jan. 21, 1941; Dept. of Biology, Sacramento
State College, Sacramento, Calif.
Education--B.A., Univ. of Texas, Austin, 19~3; M.A., 1965; Ph.D., 196~7.
Academic Experi~lzce-Assistant Prof. of Biology, Sacramento State College, 1967-
Dissertatio·n-"Taxon~m;ic, Morphological and Physiological Studies of the Genera Nostoc
and Al~abaena."
Interests--Teaching, introductory biology, plant morphology, and phycology; research,
taxonomy and morphology of the Cyanophyta.

KAPRAUN, DONALD FREDERICK-b. Springvalley, Ill., Sept. 13, 1945; Dept. of Biology, Univ. of
Southwestern La., Lafayette, La.
Education--B.S., Eastern Ill. Univ., 1966; Ph.D., Univ. of Texas, Austin, 1969.
Academic ExpeTience-Assistant Prof. of Botany, Univ. at Southwestern La., 1969-
Honors and AwarcEs--NSF Scholarship, Va. Institute of Marine Science, summer, 1964.
Dissertntion--"Field and Cultural Studies of the Genera Ulua and Enteromorpha.''
Interests-Teaching, general biology, general botany, phycooigy; research, experimental
phycology and marine ecolo·gy.

LANG, NORMA J.-b. Memphis, Tenn., July 25, 1931; Dept. of B·otany, Univ. of Calif., I)avis,
Education--B.S., Ohio State Univ., 1952; M.A., 1959; Ph.D., Ind. Univ., 1962; NIH Post-
doctorate (with H. C. Bold and W. G. Whaley), Univ. of Texas, Austin, 1962-63.
Academic Experience-Assistant Prof. of Botany, Univ. of Calif., Davis, Calif., 1963-69;
Associate Prof., 1969-
Honors and Awards-NII-I Post-doctoral Research Fellowship, Austin, Texas, 1962-63;
Editorial Board, Jo2lmal of Phycology, 1987-  ; Guggenheim Memorial Foundations Fellow-
ship, 1969-70; Darbaker Award in Phycology, 1969.
Interests--Electron microscopy of blue-green algae and Volvocales, with emphasis on sub-
cellular development and differentiation.

MAHER, SISTER MARIA LAWRENCE-b. Susq. County, July 16, 1897; Dept. of Biology, Mary-
wood College, Scranton, Pa.
Educct·tion--B.S., Marywood College, 1928; M.A., Villanova Univ., 1940; Ph.D., Fordham
Univ., 1946.
Academic Experielzce-Assistant Prof. of Biology, Marywood College, 1945-50; Associate
Prof., 1950-56; Prof., 195~-
Dissertation--"Role of Certain Environmental Factors in Growth and Reproduction of
Protosiphon botryo·ides-~lebs.'

MATTOX, KARL R.-b. Cincinnati, Ohio, Aug. 22, 193tj; Dept. of Botany, Miami Univ., Oxford,
Ohio .
Education--B.S., Miami Univ., 1958; M.A., 19~0; Ph.T>., Univ. of Texas, Austin, 196~2.
AcacE~mic Experience-Assistant Prof. of Botany, Univ. of Toronto, 1962-66; Assistant
Prof. of Botany, Miami Univ., 19BG
Dbsertation--"The Taxonomy of Certain Ulotrichalean Algae."
Interests--Teaching, fresh-water phycology, plant morphology; research, developmental
morphology of Ulotrichales, ecology of soil algae.

MCLEAN, ROBERT JAMES-b. New Haven, Conn., Aug. 15, 1940; Dept. of Biological Sciences,
State Univ. College at Brockport, Brockport, N. Y.
Education  R.A., Univ. of Conn.; M.S.; Ph.D., Univ, of Conn., 1967; N.I.H. Post-doctoral
(with H. C. Bold), 1967-68.
Academic Experie7lce-Assistant Prof. of Biology, State Univ. College at Brockport, 196~
Honors and Awards--NASA Pre-doctoral Fellowship, 19~3-~; Traineeship for Botany
Course, Woods Hole, Mass., 1964; Conn. State Graduate Scholarship, 196~-67; N.I.H. Post-
doctoral Fellowship, 1967-68.
Intetests--Algal fine structure, pigmentation, physiology of resistant algal cells.

METZNER, JEROME-b. New York City, N. Y., April 14, 1911; Dept. of Biology, John Jay
College City Univ., New York, N. Y.
Education--B.A., City College of N.Y., 1932; M.A., Columbia Univ., 1933; Ph.D., 1914.
Academic ExpeTience-Instruct~·r of Biology, Bronx High School of Science, 1944-50;
Chairman, I~pt. of B·iology, New k'ork City High Schools, 1950-~7; Prd. of Biology, John Jay
College City Univ., 1967-
Honors and Awards--Sigma Xi, Science Expert and Administrative Officer of the U. S.
Dept. o~ Army Educational Commission to Korea.
Dissertation--"A Morphological and Cytdogical Study of a New Form of Volvox."
Intel·ests-Teaching, cell biology, human biology; research, cell biology, pmtozoology, phy-

MILLER, MARCIA MADSEN-b. Santa h/lonica, Calif., May 31, 1944; Graduate Student, Dept. of
Botany, UCLA.
EcEucQtion--B.S., Univ. of Calif., Davis, Calif., 1965; M.A., 1986; Pre-doctoral (one se-
mester, with H. C. Bold), Univ. of Texas, Austin, 1966.
Academic Erperience--Graduate student at Dept. of Botany, UCLA.
Honors and Awards--Turtox, Croasdale Scholarship to Woods Hole, summer of 1985.
Interests--Teaching, cell physiology; research, chloroplast development and photosyn-

MILLIGER, LRRRY EDWARD-b. Springfield, Mo., Jan. 26, 1941; Dept. 04 Biology, Southwest
Mo. College, Springfield, Mo.
Education--B.A., Univ. of Texas, Austin, 1963; M.A. (with H. C. Bold), 1965; Ph.D., North
Texas State Univ., 1969.
Academic Experience--Curator of algal cultures, Ind. Univ., 1965-66; Assistant Prof. of
Biology, Soutl~west Mo. State College, 1969-
Honors and Awards--NSF Research Fellowship, North Texas State Univ., 1965; Sigma Xi.
Thesis--"Some Soil Algae from Bastrop State Park, Texas."
Interests--Dispersal of viable microorganisms by aquatic insects; measurement of produc-
tivity in aquatic ecosystems; the effects of pollution on ecology of inland and est~arine waters;
ecology, physiology, and distribution of fresh-water and soil algae.

MONROE, JAMES H.-b. Marietta, Okla., Aug. m, 1938; Dept. of Viral Biophysics, John L.
Smith Memorial for Cancer Research, Charles Pfizer & Co., Inc., Maywood, N. J.
Education--B.A., Univ. of Texas, 1959; Ph.D., Univ. of Texas, Austin, 1965.
Academic Experience--Research Scientist, Dept. of Viral Biophysics; John L. Smith Me-
morial for Cancer Reseach, Charles Pfizer & Co.
DISSERTATION-((SOIII Aspects of Morphogenesis and Ultrastructure in Funaria."
Interests--Viral oncology and ultrastructural pathology of tumors.

MULLAHY, REV. J. A.-Dept. of Biology, Loyola Univ., New Orleans, La. o.i.n.a.

OUTLAND, RODERICI; H.-Northwestern State College, Natchitoches, La. o.i.n.a.


PARKER, BRUCE COVELL-b. Rockingham, Vernlont, June 20, 1933; Dept. of Biology, Virginia
Polytechnic Institute and State University, Blacksburg, Va.
EducQtion--B.S. (Gum laude), Tufts Univ., 1955; M.S., Yale Univ., 1957; Ph.D., Univ. of
Texas, Austin, 19~0.
Academic Experience--Visiting Biologist, Brookhaven National Laboratory, summer, 1960;
Assistant Prof. of Botany, Univ. of Calif., Los Angeles, 1961-65; Associate Prof. of Botany,
Washington Univ., 1965-~i9; Prof. of B~atany, Va. Polytechnic Institute, 1969-
Honors and Awards--NSF Post-doctoral Fellow, Univ. College London, 1960-61; Secretary-
Treasurer, Phycological Society of America, 1965-87; Vice President, 1968; President, 1969;
Darbaker Prize in Phycology, 1970.
Disse7tation-"Environmenital Investigations of the Relationships of Certain So41 Algae with
Their Associated Microorganisms."
Interests-Experimental phycology, microbial ecology, phylogeny, and evolution.

PESSONEY, GEORGE FRANCIS III-b. Palestine, Texas, April 8, 1939; Dept. of Biology, Univ. of
Southern Miss., Hattiesburg, Miss.
Education--B.S., Sam Houston State College, 1981; M.A., 1963; Ph.D., Univ. of Texas,
Austin, 1968~
Academic ExpeTience-Instructor, Angleton Independent School System, 1483-65; As-
sistant Prof. of Biology, Univ. of Southern Miss., 1968-
Honors and Awards--Dow Chemical Co. Scholarship, 1964; Sigma Xi, N.I.H. Traineeship
in Environmental Health and Engineering, 1986.
Dissertation--"Field and Laboratory Investigation of Zygnemataceous Algae."

RIDGWAY, JOHN EDWARD JR.-b. Spangler, Pa., March 1, 1940; Dept. of Biology, Washington
Univ., St. Louis, Mo.
Education--B.A., Univ. of Texas, Austin, 1961; Ph.D., 1965.
Academic Experience-Assistant Prof. of Botany, Washington Univ., 1965-  ; Curator of
Bryophytes, Mo. Botanical Garden, 1965-
Honors and Awards Dresident, Central State Electron Microscope Society, 1968-69.
Dissertation--Soome Aspects of Morphogenesis, Biotic Coaction, and Ultrastructure in the
Genus Anthoceros."
Interests-Environm~ental, cytological, and ultrastructural aspects of morphogenesis in
cryptogams with emphasis on gam~togenesis, sporolgenesis and fertilization.

SAWA, TAKASHI-b. Osaka, Japan, Aug. 4, 1929; Dept. of Botany, Univ. of Toronto, Toronto,
Ontario, Canada.
Educati~rt--B.S., Osaka Univ., 1955; Ph.D., Univ. of Louisville, 1965; Post-doctoral re-
search associate (with H. C. Bold), Univ. of Texas, Austin, 1965-g6.
Academic Experience-Assistant Prof. of Biology, Sam Houston State College, 19~6-88;
Assistant Prof. of Botany, Univ. of Toronto, 196&
Honors and Awards-Post-doctoral Research Associate, Univ. of Texas, Austin, 19~5-66.
Interests--Teaching, phycology and cytology; research, taxonomy, cytology and paleontol-
ogy of the Charophyta.

SHEN, EUGENE Y. F.-b. Soochow, China, March 20, 1915; Dept. of Botany, Univ. of Texas,
Austin, Texas.
Education--B.S., Soochow Univ., China, 1936; M.S., 1940; M. A., Ind. Univ., 1953; Ph.D.,
Univ. of Texas, Austin, 1966.
Academic ExpeTience-Assistant Pro. to Prof. and Head of Botany Dept, National Taiwan
Univ., 1948-65; Associate Prof. of Bidlogy, Tarleton State College, 196~-67; Associate Prof. of
Biology, Wisconsin State Univ., 1967-88; research scientist, Univ. of Texas, Austin, 196~j-
Honors and Awards--Sigma Xi, Fulbright Scholarship, 1951-53 and 19612~64.
Dissertatiorl~-"Morphogenetic 'and Cytological `Investigation of Chara contraria and C. zey-
lan iccl."
Interests--Phycology and cell biology.
Contributions in Phycology

SINGH, KAMALA PRASOD-b. Girdkot, Allahabad, India, July 20, 1925; Albany Villa, Naini Tal,
UP. India.
Education--B.Sc., Banaras Hindu Univ., 1947; M.Sc., 1949; Pre-doctoral education (with
H. C. Bold), 1952r55.
Academic Experience-Assistant Prof. of Biology (o.i.n.a. ).
Honors and Awards--Indian Phycological Society.
DissertQtion-"Studies in the Genus Trachelomonas Ehrb."
Interests-Teaching, phycology; research, cytotaxonomy of the Charophyta.

SMITH, RICHARD LAWRENCE-b. Los Angeles, Calif., March 14, 1939; Dept. of Botany, Eastern
Ill. Univ., Charleston, Ill.
Education--B.A., Univ. of Calif., Davis, Calif. (Cum laude), 1961; Licensed Medical
Technologist, traineeship, St. Joseph Hospital, Burbank, Calif., 1962; Ph.D., Univ. of Texas,
Austin, 1965.
Academic Erperience-Assistant Prof. of Botany, Eastern Ill. Univ., 1965-
Honors and Awards-Sigma Xi, NDEA Pre-doctoral Fellow.
Dissertation-"Morphological, Taxonomic and Physiological Investigations of the Algal
Genera EremosphaeTa and Oocystis."
Interest-  Teaching, phycology, microbiology; research, nitrogen metabolism in algae,
comparative nutritional studies with algae, algal flora of bark, soil algae of east-central Ill.

STARR, RICHARD CAWTHON-b. Greensboro, Georgia, Aug. 24, 1924; Dept. of Botany, Ind. Univ.,
Bloomington, Ind.
Eclucation--B.S., Ga. Southern College, 1944; M.A., Peabody College, 1947; Ph.D., Van-
derbilt Univ., 1952.
Academic Experience-Instructor in Botany, Ind. Univ., 195253; Assistant Prof., 1953-57;
Associate Prof., 1957-fSO; Prof., 1960-  ; instructor in Marine Botany course at Woods Hole,
Mass., summer, 1952-55, 57-58; in charge of course 1959L63.
Honors and Aioards--l)arbaker Prize, 1955; Fulbright Fellow (pre-doctoral), Cambridge
Univ., 195~51; Guggenheim Fellow, 1959; Botanical Society of America, Secretary, 1984-69;
Phycological Society of America, President, Vice President, Secretary and Treasurer; Intema-
tional Phycological Society, Secretary, 1964-88.
Interests--Director of the culture collection ofalgae at Ind. Univ.; cultivation of the algae
with special interest in colonial Volvocales; cellular differentiation in Volvox.

SUN, ELAINA, SI-CHIN LIU-b. Si-an, China, Dec. 24, 1939; Western Michigan Univ., Kalamazoo,
Education--B.S., Taiwan National Univ., 1963; M.A., Univ. of Texas, Austin, 1966.
Academic Experience--Graduate student, Dept. of Botany, Purdue Univ., 1966-67; re-
search aid, Dept. of Biology, Western Mich. Univ., 1970-
Honors and Awards--Sigma Delta Epsjlon.
Thesis--"Studies of East Texas Soil Algae with Special Reference to Botrydiunz."

THOMAS, RUTH B.-Dept. of Biology, Sam Houston State College, Huntsville, Texas.
DtsseTtation-"Reproduction in Pinus uirginiana Miller."

THOMAS, DEMPSEY LEE-b. Macclenny, Fla., Feb. 2, 1933; Dept. of Biology, La. State Univ. at
New Orleans, New Orleans, La.
Education--B.S.A., Univ. of Fla., 1955; M.Ag., Univ. of Fla., 1958; Ph.D., Univ. of Texas,
Austin, 1968 (with R. M. Brown).
AccFdemic E 2Cp erience--Ins tructor of Biology, Saratosa High School, 195~64; Biology Con-
sultant, Univ. of Madras, India, 1964; Assistant Prof. of Biology, Concordia College, Austin,
Texas, 1965-67; Assistant Prof. of Biology, La. State Univ. at New Orleans, 1968-
Dissertation-"Morphological and Biochemical Taxonomy of the Algae Genus Protosiphon."
Interests-Morphology and biochemical systematics of algae, plant physiology.
Parker and Brown

TRAINOR, FRANCIS R.-b. Pawtucket, R. I., Feb. 11, 1929; Dept. of Biology, Univ. of Conn.,
Storrs, Conn.
Education--B.S., Providence College, 1950; M.A., Vanderbift Univ., 1953; Ph.D., 1957.
Academic Experience-Instructor of Biology, Univ. of Conn., 1957-60; Assistant Prof.,
1960-64; Associate Prof., 1964-67; Prof., 19ti7-
Honors arLd Awards-Distinguished Faculty Award in Teaching, Univ. of Conn., 196~2;
Darbaker Award, 1965; Phycological Society of America, Vice President, 196'1; President, 1968.
Dissertation--"A Comparative Study of Sexual Reproduction in Four Species of ChlQmy-
~Interests-Nutrition, structure, and sexuality in green algae and diatoms.

UHLIK, DAVID JAMES-b. Cleveland, Ohio, June 26, 1945; graduate student, Univ. of Mo.,
Columbia, Mo.
Education--B.S., St. Edwards Univ., 1987; M.A., Univ. of Texas, Austin, 19~9.
Academic Experience--Graduate Student at Univ. of Mo., currently working toward a
Ph.D. on the genetics of Chlamydomonas under Dr. C. S. Gowans.
Thesis--"Studies in ChlamzJdomonas; A. Mutagenesis in C. chEamydogama, B. Two New
Species of CiLlamydomonas."
Interests--Genetics of the fiagellar apparatus in ChlamrJdomonas eugametos.

VOSWINKEL, LINDA LOUISE-b. NOV. 1, 1944, Houston, Texas; Dept. of Environmental Engineer-
ing and Civil Engineering, Texas A&M Univ., College Station, Texas.
Educatio·n-B.A., Sam Houston State College, 19~6~; M.S., Univ. of Texas, 1969 (with R.
M. Brown).
Academic Experience--Research technician, Dept. of Environmental and Civil Engineering,
Texas A&M Univ., 1968-
Thesis--'"raxonomy and Ultrastructure of the Algal Genus Pyrobotrys."
Intel~'ests-Isolation and culturing of algae from waste stabilization ponds.

WALNE, PATHICIA L.-b. Newark, N. J., Nov. 27, 1932; Dept. of Botany, Univ. of Tenn., Knox-
ville, Tenn.
Education--B.S., Hanover College, 1954; M.S., Ind. Univ., 19S9; PbD., Univ. of Texas,
Austin, 1965.
Academic Experience-Post-doctoral research associate, Cell Research Institute and Dept.
of Botany, Univ. of Texas, 1965-68; Assistant Prof. of Botany, Univ. of Tenn., 1966-69;
Associate Prof. of Botany, 1969-  ; consultant, Biology Division, Oak Ridge National Labora-
Honors and Awards--Sigma Xi, Phi Kappa Phi; N.T.H. Pre-doctoral Fellowship, NSF Sum-
mer Fellowship; Univ. of Texas Graduate Fellowship; Turtox Croasdale Fellowship in Marine
Botany; Post-doctoral Research Associate, Univ. of Texas, Austin; Editor, Phycological News-
Dissertation-"Cellular Organization in Chlamydomonas and the Effects of Colchicine."
Interests-- Teaching, phycology, cell biology, experimental cytology; research, development
and characterization of eye spots in euglenoid and chlamydomonad flagellates, ultrastructural
physiological and serological study of developments in euglenoid and chlamydomonad flagel-

WATSON, MELVIN WILLIAM-b. Louisville, Ky., Sept. 29, 1949, graduate student; Dept. of
Botany, Univ. of Texas, Austin, Texas.
Education--B.S., Univ. of Louisville, 19Ci4; M.S., 1967; Ph.D., Univ. of Texas, Austin, 1970
(with H. G. Arnott).
Dissertation-"Ultrastructural Aspects of Development in Microthamnion."
In~erests-Experimental phycology, especially ultrastructure and development of syn-
chronous cultures.

WIEDEMAN, VARLEY-Dept. Of Biology, Univ. of Louisville, Ky. o.i.n.a.


This is the alga which Walter Herndon discovered and named after Harold

Photo of Harold C. Bold which appeared in our Book

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